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The goal of the Howard Hughes Medical Institute (HHMI) New Generation Program is to develop an entire generation of new resources to enhance undergraduate biology education. Our work is guided by the following hypothesis: Training graduate students to teach and mentor improves the education of undergraduates and the preparation of graduate students for careers in science. We promote graduate students and postdocs as vital educational resources by providing them the education necessary to become outstanding mentors and classroom teachers. Specifically, graduate students and post-docs are invited to apply for HHMI Teaching Fellowships that support activities directed toward building their skills as teachers and mentors. Each teaching fellow takes “Teaching Biology,” a course that covers theoretical and practical aspects of biology education, providing students with a structured setting in which to practice and receive feedback on teaching. During subsequent semesters, each fellow chooses to serve as a mentor for an undergraduate researcher, teach in a classroom, or develop instructional materials. Each of these initiatives will be described. In addition, we are studying the impact of this training comprehensively, including examination of the participating students and postdocs, the undergraduates they teach and the faculty advisors with whom they work.